Origins: How Customisable are related empires?

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In Scion, you will start as a vassal of either the spiritualist or materialist Fallen Empires. If you're spiritualist or materialist, it will pick the one that mirrors your ethic. Otherwise it's randomized. For the Origins where you start in a federation, the other two members are randomized but their ethics will largely reflect those of your empire.
 
For Lost Colony, it's completely random. They could be anything from purifiers to pacifists. It's been a while since your people left the homeworld - a lot will have happened in that time.
 
The other two members of your federation will also start with the Common Ground/Hegemon origin (called Hegemon Subordinate in the case of that origin).

Alas, it will not be possible to start with human players as the other federation members in multiplayer.