Going to the religion tab, the holy sites portraits are all black squares, with a random 3 coa, with the flag green and star on, despite not owning one of the sites. See picture for a better idea. Loaded the save that was started in 2.1.4, and was played in all the beta patches. Last saved while playing in 2.1.5.4, which did not have this problem last time. Only popped up now, after loading from a CTD after checking a dead character's page. Also, notice the straighten up button still grayed out due to being asked from someone else.
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